Delectopecten zacae (Hertlein, 1935)
HERTLEIN, L. G. 1935. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. The recent Pectinidae. Proceedings of the California Academy of Sciences [4th Series], 21 (25): 301-328, pls. 18-19. [p. 321, pl. 18, figs. 3-6, 9-10]
1908 Pecten (Pseudamusium) [sic] panamensis Dall, 1908
1935 Pecten (Delectopecten) zacae Hertlein, 1935
1935 Pecten (Delectopecten) zacae Hertlein, 1935
L. G. Hertlein, 1935, plate 18.
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«Pecten (Pseudamusium) panamensis Dall, Bull. Mus. Comp. Zool., vol. 43, no. 6, 1908, p. 404, pl. 6, figs.8 and 10. "Gulf of Panama, in 322 fathoms, mud, bottom temperature 56ºF."
Not Pecten panamensis Dall, Trans. Wagner Free Inst. Sci., vol. 3, pt. 4, 1898, p. 696. No locality cited [? :P. fasciculatus Hinds]. Range: Cape San Lucas, Lower California to Panama. Holotype: No. 6880 (C. A. S. Paleo. type collection), from Loc. 27587 (C. A. S.), off Cape San Lucas, Lower California; paratypes: Nos. 6881, 6882 (C. A. S. Paleo. type collection), from Loc. 27584 (C. A. S.), dredged in 20 to 220 fathoms from Lat. 23° 03' to 23º 06' N., Long. 109° 31' to 109° 36' W.; Templeton Crocker collector. This species was dredged by the Templeton Crocker Expedition at two localities. Loc.27584 (C. A. S.), about 10 miles due east of San Jose del Cabo, Lower California, in 20 to 220 falhons, Lat. 23° 03' to 23º 06' N., Long. 109° 31' to 109° 36' W.; and. Loc. 27587 (C. A.S.), oft Cape San Lucas, Lower California, in 20 to 25 fathoms. The thin, delicate forms of this species, collected by the Templeton Crocker Expedition, possess the shell characters of the species described by Dall as P. panamensis in 1908 (not P. panamensis DaIl, 1898). Due to the fact that Dall has used the name panamensis for a species in 1898, the name zacae is here proposed for the present species. It is named for Mr. Templeton Crocker's yacht, Zaca. Some specimens are nearly smooth, while others have 40 to 65 fine radial, minutely scaly threads. The whole surface externally, is covered by fine camptonectes striations. The more elongate form, fewer and more widely spaced ribs, the broader anterior ear of the right valve, and the squarer posterior margin of the posterior ears of both valves, separate the species from P. randolphi Dall, and P. randolphi tillamookensis Arnold. Pecten lillisi Hertlein [53] from the Kreyenhagen Shale (upper Eocene or lower Oligocene), appears to be a related species.» 53 Pecten (Pseudamusium) lillisi Hertlein, Bull. South Calif. Acad. Sci., vol. 33, pt. 1, January-April (issued February 28), 1934, p. 5, pl. 1, fig. 1, pl. 2, figs. 2, 3. "diatomite, Kreyenhagen shale, from S. E. corner of Sec. 35, T. 6 S., R. 7 E., M. D. M., Stanislaus County, California; on the oorth side of Crow Creek road." LEO GEORGE HERTLEIN, 1935
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«Addiitional descriptive notes: Specimens from the Gulf of California are more convex, smoother and glossier than those from the Gulf of Panama, and often have a few very faint concentric undulations of the disks below the umbonal region. Dall was correct in surmising that the "minute beads on the lower margin of the fasciole" comprise a ctenolium in the juvenile stage; the teeth become bead-like after the shell reaches an altitude of from 10 to 14 mm. The valves seldom have similar sculpture, usually only one having pronounced radial threads.
Remarks; Dall first used the name Pecten panamensis in 1898 (loc. cit.). In discussing the "section" Pallium he said, "Pecten panamensis Dall, which has in most respects an unusually close resemblance to P. plica, differs in having the cardinal laminae obsolete and in the presence of a byssal sinus and ctenolium." Discussing genera, he said, "Bucquoy, Dautzenberg, and Dollfus (1889) propose Peplum, with P. clavatus Poli as type, which would include such species as P. panamensis." No further description (in the accepted sense of the word) was given, nor was the species figured, but the features he mentioned in the first sentence, those he implied in the second, and the locality inferred by the name, add up to what may be construed as a "definition," in accordance with Article 25 of the International Rules; thus Pecten panamensis (1898) was established as a published name. Hinds, however, had described the same species in 1845 as Pecten fasciculatus (see Pseudamussium (Peplum) fasciculatum (Hinds), this paper); of that there can be no doubt, for no other species of Peplum is found in the eastern Pacific. Therefore, Dall's Pecten panamensis (1898) was a synonym for P. fasciculatus Hinds from the time it was published. Apparently forgetting that he had previously named the species, Dall described and figured it in 1908 as Pecten (Pallium) miser, thus adding another synonym for P. fasciculatus. In the same 1908 paper Dall described and figured Pecten (Pseudamusium) panamensis; since that name had been published earlier, P. panamensis (1908) was a homonym, and Hertlein rectified the matter in 1935 by re-naming the species Pecten (Delectopecten) zacae. Geographical range: Punta San Fermin, northern Gulf of California to Panama Bay. Also Galapagos Islands. The author determined the Clipperton Islands specimens referred to this species to be Cyclopecten (Delectopecten) vitreus (Gmelin). The previously recorded northern limit for C. zacae was San Jose del Cabo, eastern Lower California (Hertlein, 1935. p. 321), but in July, 1958, Dr. H. A. Rehder, of the U. S. National Museum, sent the author 139 lots of eastern Pacific material from the Museum collection for identification, and one lot contained a specimen from Albatross station 3035, off Punta San Fermin; that record extends the northern range by about 570 miles. Geochronological range: Recorded only from the Recent. Bathymetric range: Recorded in 5 to 1005 fathoms. Ecological data: Found only on mud bottoms.» GRAU, G. 1959. Pectinidae of the eastern Pacific. Allan Hancock Pacific Expeditions, 23: i-viii, 1-308, 57 pls. University of Southern California Press. Los Angeles, California. [p. 51, 52]
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Cyclopecten (Delectopecten) zacae (Hertlein) 1935; G. Grau, 1959, Pectinidae of the eastern Pacific, plate 19.
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