Notochlamys hexactes (Péron in Lamarck, 1819)
LAMARCK,
J. B. 1819. Histoire naturelle des animaux sans vertèbres, présentant les
caractères généraux et particuliers de ces animaux, leur distribution, leurs
classes, leurs familles, leurs genres, et la citation des principales espèces
qui s'y rapportent. Tome sixième, Premier partie, 343 p. Chez l'Auteur, au
Jardin du Roi. De l'imprimerie de A. Belin.
Paris, 1819.
[p. 178, sp. 59]
1819 Pecten hexactes Péron in Lamarck, 1819
1842 Pecten undulatus G. B. Sowerby II, 1842
1852 Pecten pesanatis Reeve, 1852
1864 Pecten tasmanicus Adams & Angas, 1864
1876 Pecten mariae Tenison Woods, 1876
1927 Chlamys anguineus Finlay, 1927 [nomen novum pro Pecten undulatus G. B. Sowerby II, 1842]
1842 Pecten undulatus G. B. Sowerby II, 1842
1852 Pecten pesanatis Reeve, 1852
1864 Pecten tasmanicus Adams & Angas, 1864
1876 Pecten mariae Tenison Woods, 1876
1927 Chlamys anguineus Finlay, 1927 [nomen novum pro Pecten undulatus G. B. Sowerby II, 1842]
Notochlamys hexactes (Péron in Lamarck, 1819); B. K. raines & G. T. Poppe, 2006, A Conchological Iconography, The Family Pectinidae, pl. 169.
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«59. Peigne à-six-rayons. Pecten hexactes.
P. testâ albâ, flabellatâ; radiis sex longitudinaliter striatis; medianis latioribus.
Pecten hexactes. Péron. Habite les mers de la Nouvelle Hollande, au port du Roi Georges. Mus. nº, Largeur, 35 millimètres. Etc.» JEAN BAPTISTE LAMARCK, 1819
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«Remarks. Notochlamys hexactes inhabits only the temperate waters of southern Australia and is extremely variable in shape (left valve flat and right valve weakly convex to equally inflated and pyxoid), sculpture (arrangement, number and prominence of primary radial plicae and secondary radial ribs) and colour (typically highly variable and brightly coloured, Tasmanian morph usually only purple). The “western” (typical or anguinea morph) and “eastern” populations (tasmanica morph) intergrade completely and are not distinguishable geographical forms (see also Beu & Darragh, 2001: 66). As noted above under Equichlamys bifrons, Notochlamys hexactes has lecithotropic development, and the great range of variation presumably results from the poor exchange of genes between distant populations.»
DIJKSTRA, H. H. & A. G. BEU. 2018. Living scallops of Australia and adjacent waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae). Records of the Australian Museum, 70 (2): 113-330, figs. 1-102. [p. 251]
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Notochlamys hexactes (Lamarck); H. H. Dijkstra & A. G. Beu, 2018, Living scallops of australia and adjacent waters, figures 74A-74D.
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«Type material. Holotype of Pecten hexactes Lamarck, 1819, MNHN (Dijkstra 1995: pl. 30, figs 130, 131), from King George Sound, southern Western Australia. Pecten undulatus Sowerby, 1842, 5 syntypes BMNH 1951.3.28.4- 5 (3 LVs and 2 RVs; Dr T. R. Waller 1999, pers. comm.); Sowerby (1842: 60) did not know the provenance of the type material, and suggested ‘Mediterranean?’, and it still needs to be clarified whether this nominal species actually is conspecific with N. hexactes. (Which introduces the possibility that the type species of Notochlamys was misidentified.) Dr T. R. Waller has informed us that the LV illustrated by Reeve (1852: pi. 20, fig. 73) is among these syntypes, but that illustrated by Sowerby (1842) is not present. Holotype of Pecten tasmanicus A. Adams & Angas, 1864, not seen, BMNH 1950.5.23.1 (H. H. Dijkstra, pers. comm.), from Oyster Bay, Tasmania; type material of Pecten mariae Tenison Woods, 1876, not seen, Tasmanian Museum TM 5749 (LV) (H. H. Dijkstra, pers. comm.) from ‘East Coast and Maria Island’, Tasmania.
Occurrence and time range. We know of no pre-Holocene fossil records. At present, N. hexactes ranges from southern Western Australia (AMS, 16 lots, northernmost Cottesloe Beach, Fremantle), South Australia (7 lots), Victoria (5 lots), and Tasmania (6 lots) to southern New South Wales (1 lot, the specimen recorded from Twofold Bay by Iredale 1924). Holocene fossils are present in WAM from Fremantle, lower Swan Estuary, East Rockingham, Cockburn Sound and Albany, all in southern Western Australia. The fossil record suggests that it may be a very recent arrival in southwestern Western Australia. Remarks. Examination of a large range of material in AMS (35 lots) showed that a single, highly variable species of Notochlamys inhabits southern Australia, ranging (like most of the other endemic southern Australian scallops) from southern Western Australia to southern New South Wales, including all of Tasmania. The population is very variable in the width, number and prominence of the plicae, as well as in the number and coarseness of the radial costae and in shell color. Although specimens with numerous, narrow plicae and many relatively prominent costae (the tasmanica form) tend to predominate in the south and east of its range, and specimens with relatively few, prominent, wide plicae and numerous weak radial costae (the anguinea form) tend to predominate in southern Western Australia, the two forms intergrade completely, the same variants occur throughout the range, and there is no doubt that this is a single, highly variable species. Specimens are illustrated here to compare with the fossil species referred to Notochlamys, and to determine the characters of the LV preradial dissoconch sculpture. Dijkstra (1995: 492, pi. 30, figs 130, 131) has shown that Lamarck’s (1819: 178) holotype of Pecten hexactes is a small, very coarsely plicate RV of the species usually known as Notochlamys anguinea, with six plicae, collected in King George Sound, southern Western Australia, by Péron.» BEU, A. G. & T. A. DARRAGH. 2001. Revision of southern Australian Cenozoic fossil Pectinidae (Mollusca: Bivalvia). Proceedings of the Royal Society of Victoria, 113: 1-205. [p. 65, 66]
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Notochlamys hexactes (Lamarck); A. G. Beu & T. A. Darragh, 2001, Revision of southern Australian Cenozoic fossil Pectinidae, figure 17.
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«REMARKS:
Péron's handwritten label: "Port du Roi George. Pecten Hexactes" is glued on Lamarck's label: "peigne à-six-rayons. p. hexactes" on the back of a cardboard plate. There was no description published for this species, as it was a manuscript name. During the visit to King George Sound (see above), Péron collected many shells along the shore, as reported in Baudin's journal: " ... He has one or two cases of broken shells, for in several places along the shore one can shovel them up...." (CORNELL, 1974 : 494). CURRENT TAXONOMIC POSITION AND DISTRIBUTION: P. hexactes Péron in LAMARCK is a senior synonym of Pecten tasmanicus Adams & Angas, 1863 (= Chlamys anguineus Finlay, 1927), and actually provisionally placed in Notochlamys Cotton, 1930 (under study). N. hexactes (Péron in LAMARCK) is distributed along the coast of southern Australia from south Western Australia to Victoria and Tasmania. Living specimens are littoral to sublittoral, and are observed among rubble in silty reefs or under rocks on sandy bottoms.» DIJKSTRA, H. H. 1994. Type specimens of recent species of Pectinidae described by Lamarck (1819), preserved in the Muséum d'Histoire Naturelle of Geneva and the Muséum National d'Histoire Naturelle of Paris. Revue Suisse de Zoologie, 101 (2): 465-532, 30 pls. [p. 493]
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Pecten hexactes [Péron] Lamarck, 1819 H. H. Dijkstra, 1994, Type specimens of recent species of Pectinidae described by Lamarck, plate 30, figures 130-132
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