Complicachlamys wardiana Iredale, 1939
IREDALE, T. 1939. Mollusca. Part 1. In Great Barrier Reef Expedition 1928-29, B.M.(N.H.), Scientific Reports, 5 (6): 209-425, pls. 1-7. [p. 362, pl. 5, figs. 25, 25a]
T. Iredale, 1939, plate 5.
|
«There is great difficulty in this little group, as Melvill and Standen recorded crouchi and dringi as separate species from Queensland. Odhner included fulvicostatus and dringi as separate species from North-West Australia. Lynge (p. 156) and Bavay synonymized dringi withfulvicostatus, and added luculentus.
Pecten crouchi was described by Smith ('AIUl. Mag. Nat. Hist.' IX, ser. vi, p. 255, fig. in text, March, 1892) from Mauritius with very unequal auricles and nine ribs, each bearing seven raised lines. The name can be dismissed at once from the Australian list, as crouchi can only be regarded as a geographical representative of dringi, and therefore could not occur in Queensland. Pecten dringi was introduced by Reeve forty years previously ('Conch. Icon.' VIII, pI. xxxiii, sp. and fig. 152, August, 1853) for a species collected by Dring at Bathurst Island, north-west coast of Australia. The species, which Odhner added from northwest Australia, was described by Adams and Reeve (Pecten fulvicostatus, 'ZooI. Voy. "Samarang", Mollusca', p. 74, pI. xxi, fig. Il, 1850) from the Sooloo Archipelago, and the figure in shape and ribbing does not agree with Australian specimens. Bavay's addition of luculentus is correct, as Reeve ('Conch. Icon.' VIII, pI. xvi, sp. and fig. 59, February, 1853) had proposed Pecten luculenta from Bathurst Island, North Australia, and apparently overlooked it, when a few months later he named dringi from the same locality. In connection with the latter Reeve remarked "Colouring extremely variable", but the shape also varies and thus accounts for the attempts to record two species where only one exists. A fair series of specimens is available from Darwin, practically the type locality of luculenta and dringi, and others from North-West Australia, and while these show variation they are separable from the Queensland specimens, so the latter are named as above. The type is a specimen collected by Mr. Melbourne Ward at Hayman Island, Whitsunday Passage, measuring 29'5 mm. long by 25 mm. broad. A series of luculenta = dringi from Broome, North-West Australia, ranges up to 60 mm. long by 50 mm. broad. A shell from the latter series measuring 29 mm. long by 25 mm. broad has been compared with the Hayman Island type, and it has the ribs much more elevated, the intervals consequently deeper and wider and more marked; this is still more noticeable in the left valve, where the median riblet of each rib has broadened and become more distinct. The Queensland shell ranges along the coast line of Queensland from Cape York to Port Curtis, an odd specimen occurring at Low Isles. It may be possible that the species here distinguished can be degraded to the rank of geographical races or subspecies, but the experience gained in the study of marine molluscs does not make this a desirable course without a very complete knowledge of the animals concerned. Therefore we may allow at present: Complicachlamys fulvicostata A. Adams and Reeve, 1850. Sooloo Archipelago. Complicachlamys luculenta Reeve, February, 1853 = dringi Reeve, August, 1853. North and North-West Australia. Complicachlamys crouchi Smith, 1892. Mauritius. Complicachlamys wardiana Iredale, 1938. Queensland.» TOM IREDALE, 1939
|
«Description. Shell up to 60 mm high, most specimens to c. 40 mm; rather solid, somewhat prosocline, left valve flattened, right valve slightly more inflated, inequivalve, inequilateral, auricles highly unequal in shape and size, umbonal angle c. 80–85°; colour highly variable, white, cream, yellowish, orange, red, purple, brown or almost black; uniform, mottled or with paler zig-zag streaks.
Both valves with 7–9 (8 on most specimens) evenly spaced, low (in Queensland morph) to more angulate (in WA morph) radial plicae. Numerous closely spaced radial riblets on plicae (3–5) and in interspaces (7–10), with rather coarse lamellae on plicae in WA morph, lamellae delicate or lacking in Queensland morph. Microscopic interstitial commarginal lamellae in WA morph, more minute or lacking in Queensland morph. Anterior auricles much larger than posterior, with c. 8–10 fine radial riblets on left valve, c. 5–6 more prominent ones on right valve. Byssal notch deep, byssal fasciole rather broad. Functional ctenolium with c. 8 well-developed teeth Dimensions. Illustrated specimens: WA, Broome, Entrance Point (AM C.057307): rv: 57.4, L 49.9 mm; lv: H 57.2, L 49.6 mm; D 12.9 mm; NT, Darwin Harbour, East Arm, 12°30'S, illustrated by Lamprell & Whitehead (1992: pl. 12, fig. 72) (AM C.303787): rv: H 45.2, L 40.1 mm; lv: H 45.2, L 38.8 mm; D 11.8 mm. Iredale (1939: 363) stated the dimensions of the holotype as H 29.5, L 25 mm.
Habitat. Living in the intertidal to sublittoral zones, byssally attached to undersides of coral slabs on hard reef at extreme low tide, often in silty conditions. Some adults become detached and lie in the silt under slabs.
Distribution. Tropical Indo-West Pacific from China to the Gulf of Siam and from the Philippines to northern Australia (Raines & Poppe, 2006: 188); Vietnam to Australia, 0–180 m (Huber, 2010: 209); Philippines, 14–90 m (Dijkstra, 2013: 58); South China Sea (as “Semipallium fulvicosta”) (Xu & Zhang, 2008: 87); Indonesia, Sulawesi (Dijkstra, 1991: 38); Maximum depth range of live-taken specimens is from the intertidal zone to 180 m. Present specimens from Australia alive in the intertidal zone to 73 m.
Remarks. Specimens from northwestern Australia have coarser sculpture than those from Queensland, but specimens from central northern Australia intergrade slightly. Examined material from the Indo-West Pacific is indistinguishable from the type material from Queensland.»
DIJKSTRA, H. H. & A. G. BEU. 2018. Living scallops of Australia and adjacent waters (Mollusca: Bivalvia: Pectinoidea: Propeamussiidae, Cyclochlamydidae and Pectinidae). Records of the Australian Museum, 70 (2): 113-330, figs. 1-102. [p. 229, 230]
|
Complicachlamys wardiana Iredale; H. H. Dijkstra & A. G. Beu, 2018, Living scallops of australia and adjacent waters, figures 59D, 59E, 59G, 59H, 59J, 59K, 61B, 61C.
|
«Type data. — Holotype (H 29.5 mm, pv) AMS C.90373. Type locality: Queensland, Whitsunday Passage, Hayman Island.
The paralectotypes of Pecten dringi Reeve, 1853 are in fact Complicachlamys wardiana Iredale, 1939, collected from Bathurst Island, see Wagner (1989b: 114). The lectotype is Pecten dringi Reeve, 1853, a species in Semipallium. Material examined. — Philippines: PMBP 2004, Stn L46, 90-110 m, 1 pv (A); Stn S10, 6-14 m, 1 pv (A).
Description. — Shell up to 60 mm high, most specimens to c. 40 mm; rather solid, somewhat prosocline, left valve flattened, right valve slightly more inflated, inequivalve, inequilateral, auricles highly unequal in shape and size, umbonal angle c. 80-85˚; colour highly variable, white, cream, yellowish, orange, red, purple, brown or almost black; uniform, mottled or with paler zig-zag streaks. Both valves with 7-9 (8 on
most specimens) evenly spaced, low (in Queensland-morph) to more angulate (in WA-morph) radial costae. Numerous closely spaced radial riblets on costae (3-5) and in interspaces (7-10), with rather coarse lamellae on costae in WA-morph, lamellae delicate or lacking in Queensland-morph. Microscopic interstitial commarginal lamellae in WA-morph, more minute or lacking in Queensland-morph. Anterior auricles much larger than posterior, with c. 8-10 fine radial riblets on left valve, c. 5-6 more prominent ones on right valve. Byssal notch deep, byssal fasciole rather broad. Functional ctenolium with c. 8 well-developed teeth. Distribution. — Tropical Indo-West Pacific from China to the Gulf of Siam and from the Philippine Islands to northern Australia (Raines & Poppe, 2006: 188). Present specimens from the Philippines alive at 14-90 m (minimum depth range). Living byssally attached to undersides of coral slabs on hard reef at extreme low tide, often in silty conditions. Adults may become detached and lie in the silt under slabs. Remarks. — Present material from the Philippines is indistinguishable from the type material from Queensland. Specimens from northwestern Australia have coarser sculpture than those from Queensland, but specimens from central northern Australia intergrade slightly.» DIJKSTRA, H. H. 2013. Pectinoidea (Bivalvia: Propeamussiidae and Pectinidae) from the Panglao region, Philippine Islands. Vita Malacologica, 10: 1-108, pls. 1-32 pls, 2 tabs. [p. 57, 58]
|
Complicachlamys wardiana Iredale, 1939; H. H. Dijkstra, 2013, Pectinoidea from the Panglao region, plate 15, figures 3a-3d.
|