Weyla alata (Buch, 1838)
BUCH, L. VON. 1838. Über den zoologischen Character der Secundär Formationen in Süd-Amerika. Bericht über die zur Bekanntmachung geeigneten Verhandlungen der Konigl. Preuss. Akademie der Wissenschaften zu Berlin, 1838, p. 54-67. [p. 55]
1838 Pecten alatus Buch, 1838
1842 Pecten dufrenoyi Orbigny, 1842
1864 Pecten acutiplicatus Meek, 1864
1899 Pecten paragogus R. A. Philippi, 1899
1899 Pecten angustecostatus R. A. Philippi, 1899
1899 Pecten subcarinatus R. A. Philippi, 1899
1899 Pecten stolpi R. A. Philippi, 1899
1899 Pecten commutatus R. A. Philippi, 1899
1914 Vola alata var. aspera Jaworski, 1914
1929 Neithea mexicana Jaworski, 1929
1933 Parapecten praecursor Crickmay, 1933
1987 Weyla (Lywea) meeki Damborenea, 1987 [nomen novum pro Pecten acutiplicatus Meek, 1864]
1842 Pecten dufrenoyi Orbigny, 1842
1864 Pecten acutiplicatus Meek, 1864
1899 Pecten paragogus R. A. Philippi, 1899
1899 Pecten angustecostatus R. A. Philippi, 1899
1899 Pecten subcarinatus R. A. Philippi, 1899
1899 Pecten stolpi R. A. Philippi, 1899
1899 Pecten commutatus R. A. Philippi, 1899
1914 Vola alata var. aspera Jaworski, 1914
1929 Neithea mexicana Jaworski, 1929
1933 Parapecten praecursor Crickmay, 1933
1987 Weyla (Lywea) meeki Damborenea, 1987 [nomen novum pro Pecten acutiplicatus Meek, 1864]
Pecten alatus; L. von Buch, 1939, Pétrifications recueillies en Amérique, figures 1-4.
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«Die gröfsere Menge besteht aus einem PECTEN. Aus der ausgezeichneten Abtheilung, welche durch den weit verbreiteten Pecten quinquecostatus so sehr bekannt Ist, und die man schon sogar als ein eigenes Geschlecht unter dem Namen NETHEA aufgeführt hat. In dieser Abthellung nehmlich ist die Unterschaale sehr hoch gewölbt, und steht mit einem bedeutenden Schnabel über die ganz flache Oberschaale hervor; eine Ungleichheit beider Schaalen, welche gar mächtig auffällt. — Diese Formen sind aber bisher noch niemals In anderen Formationen, als in der Kreide gesehen worden, selbst in America nicht, wo sie in den atlantischen Provinzen der nördlichen Freistaaten und in Alabama gar nicht selten gefunden werden. Es folgt daher hieraus die Wahrscheinlichkeit, dafs auch die Secundärformatlonen am oberen Amazonenslrome zur Kreide, und nicht zu Juraformationen gehören, und diese Wahrscheinlichkeit kann nicht eher wieder aufgegeben werden, als bis unzubezweifelnde Juraprodukte darüber eines Anderen belehren.
Der PECTEN ALATUS m. ist ungleichseitig. Die vordere Seite der gewölbten Schaale tritt an ihrem Rande hervor, und verflacht sich zu einem bedeutenden Flügel ; daher geht auch die Axe der Schaale der ungeflügelten Seite näher durch das Drlttheil der gröfsten Breite. Vierzehn hochstehende, aber ganz gleiche Falten oben nur flach gewölbt, bedecken die Schaale; ihre Zwischenräume sind fast doppelt so breit und beide sind sehr fein concentrisch gestreift. Der Rand an der Seite des Schlosses ist umgebogen, und bildet mit der, ebenfalls umgebogenen flachen Schaale eine ausgezeichnete Lunula zu beiden Seiten unter dem Schlofs, welche ohngefähr ein Vierthell der ganzen Länge der Axe einnimmt. An den umgebogenen Rändern bemerkt man eine sehr bestimmte Diagonalstreifung, welche die Anwachsstrelfen durchschneidet. Die flache Schaale ist in der Mitte etwas concav, und ist mit 12 schmalen Falten geziert, deren Breite nicht das Viertheil ihrer Zwischenräume erreichen. Die Schlofskante auf der geflügelten (Byssus) Seite ist stark ausgeschweift, oder concav, und erreicht nicht die Hälfte der Axenlänge; dagegen dehnt sich die gerade Schlofskante der gegenüberstehenden Seite bis nahe ¾ dieser Axenlänge herunter. Die Dicke der gewölbten Schaale beträgt mehr als ein Dritthell der Länge. Das Schlofs hat an dem vorderen Ohr der gewölbten Schaale einen deutlichen Byssusausschnitt, dem gemäfs die feine, senkrechte Streifung sich Im Bogen herabzieht; das hintere Ohr ist nicht ausgebogen, mit schief, dem Schnabel zu geneigtem Rande, wie gewöhnlich. Ohne die ausgezeichnete Flügelausbiegung würde der ganze Pecten dem Pecten aequicostatus sehr ähnlich sein, so wie er in der unteren Kreide, unter anderen so häufig bei Regensburg vorkommt. Dieser Pecten bildet in den peruanischen Anden, da wo er vorkommt, ganze Conglomerate, und wenn man von der grofsen Masse von Muscheln redet, die auf den Andesbergen gefunden werden, so ist er es vorzüglich, welcher dazu Veranlassung giebt und der ganze Berge zusammensetzt, wahrscheinlich auf ähnliche Art, wie ein grofser Theil von Sicilien nur allein durch den Pecten opercularis sich über die Meeresfläche erhebt. Hr. v. Humboldt hat ihn nicht allein von der Höhe von Guancavelica erhalten, sondern auch ausgezeichnet von Copiapo im 26º südl. Breite, und er selbst hat ihn in grofser Menge gefunden zwischen Guanibos und Montan in 8400 Fufs Höhe; zwischen dem Amazonenstrom und Lima. Das ist eine gewaltige Ausdehnung, und dennoch ist es sehr glaublich, dafs die ganze Formation, wenigstens bis Cuzeo hinaus, ununterbrochen fortgehe. Hr. v. Humboldt hat die Güte gehabt, mir einen Auszug aus seinen Tagebüchern zu geben , aus welchen sowohl diese Ausdehnung, als auch die geognostischen Verhältnisse dieser Massen klar hervorgehen.» CHRISTIAN LEOPOLD VON BUCH, 1838
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«Remarks: For a full description of W. (W.) alata refer to Damborenea (1987b) and Aberhan (1994). Recently, Mexican specimens of W. alata were described by Damborenea and González-León (1997). These authors concluded that Weyla mexicana (Jaworski, 1929) is a junior synonym of W. alata. Having studied Jaworski’s type material of W. mexicana and additional material from the type locality, we follow this view here. Aberhan (1998a) regarded W. mexicana as a synonym of Weyla meeki Damborenea (1987b), a problematic species first described as Pecten acutiplicatus by Meek (1864) from the Lower Jurassic of California. According to present knowledge, we cannot exclude W. meeki as another junior synonym of W. alata. Clearly, more well-preserved, articulated specimens of W. meeki are needed from the type locality to settle this issue.
In the studied sections, W. alata is by far the most common species of Weyla. In comparison with Weyla (W.) bodenbenderi (see below), its ribs are always simple and do not exhibit intercalations of secondary ribs or splitting of primary ribs. Weyla (Lywea) unca (see below) has triangular ribs on both valves and is biconvex in all ontogenetic stages.» SCHOLZ, A., M. ABERHAN & C. M. GONZÁLEZ-LEÓN. 2008. Early Jurassic bivalves of the Antimonio terrane (Sonora, NW Mexico): Taxonomy, biogeography, and paleogeographic implications. Geological Society of America, Special Papers, 442: 269-312, figs. 1-17. [p. 287]
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Weyla (Weyla) alata (von Buch, 1838); A. Scholz, M. Aberhan & C. M. González-León, 2008, Early Jurassic bivalves of the Antimonio terrane, figures 7F-7J, 7M-7O.
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«Material — Seven specimens with both valves, six left valves and eight right valves, most of them incomplete and not very well preserved (ERNO-2375, 2380 to 2387, 2399 to 2403, 2413,2414,2419,2421,2436 to 2438) from Sierra de Santa Rosa Formation, lower and middle members; Lower Jurassic (Sinemurian-Pliensbachian?); Sierra de Santa Rosa, sections 3 and 4. One right valve (ERNO-2374) from Antimonio Formation, unit 19; Sierra del Alamo.
Remarks — The specimens at hand do not show any differences from those referred to W. alata from other localities in South and North America (see Smith and Tipper, 1986; Damborenea, 1987b; Damborenea and Manceñido, 1988) and are thus referred to this species. The name W. mexicana (Jaworski) is here regarded as a junior synonym of W. alata (von Buch). It might only be maintained as a subspecific name if some distinctive morphological characters could be found. Most of the specimens here described, as well as those figured by Lucas and Estep (1997, fig. IA-lB) were collected in Sierra de Santa Rosa, i.e. the type locality of W. mexicana (Jaworski, 1929). Jaworski characterized his new species by the presence of V-shaped ribs on both valves. Nevertheless, examination of all the material from Sierra de Santa Rosa (NHMB and ERNO collections) shows that this feature, together with some peculiar surface "ornament pattern", is merely a preservational artifact. It is also interesting to note that the specimens available to Jaworski (1929) were small to medium-sized (up to 6 cm high), and Weyla shells tend to have more acute ribs at that size. The collection examined here contains specimens of a wide range of sizes and different degrees of corrosion of the shell, but clearly conspecific. Small specimens (up to 4 cm high), such as ERNO-2374, 2375, 2384, 2385, 2387, 2402, 2414 and 2421, can be closely compared with the three specimens figured by Jaworski (NHMB-51b1 , 521 and 522), but these, together with medium-sized specimens, show the same range of morphological variation as W. alata from other localities. On the other hand, very large specimens (between 12 and 16 cm high), such as ERNO-2380 (Figure 5.9) and 2399, have flat-topped ribs and a few faint radial costae on the anterior auricle of the right valve (this last characteristic is also present on smaller shells). These features, together with the large size, were used by Aberhan (1994) to characterize Weyla titan (Möricke) as a distinct species. This was described (but not figured) by Möricke (1894) as a variety of W. alata, and as such was mentioned (but again never figured) by several later authors (Philippi, 1899; Jaworski, 1914, 1915, 1925). The few known specimens of W. alata titan always appear associated with W. alata along the Andes (as in Sonora) and all their characteristic features could be due to the large size attained. It was regarded as a subspecies of W. alata by Perez and Reyes (1994), and it seems that it could be just a "gerontic" variant of W. alata. Thus Weyla specimens from Sierra de Santa Rosa are here regarded as belonging to a single species, and if deemed necessary, they could be referred to W. alata mexicana (Jaworski) (Figures 5.6-5.8) and W. alata titan (Möricke) (Figure 5.9). Weyla alata has a wide stratigraphical distribution in the Americas, from at least Sinemurian to lower Toarcian (Damborenea and Manceñido, 1988; Perez and Reyes, 1994). The material figured as Weyla by Lucas and Estep (1997, p. 45, figs. 1C-1D) from Sierra del Alamo does not belong to this species but to a coarsely ribbed limoid, which is here described as Mysidioptera mexicana n. sp.» DAMBORENEA, S. E. & C. M. GONZÁLEZ-LEÓN. 1997. Late Triassic bivalves from Sonora, Mexico. Revista Mexicana de Ciencias Geológicas, 14 (2): 178-201. [p. 190, 191]
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Weyla (Weyla) alata (von Buch, 1838); S. E. Damborenea & C. M. González-León, 1997, Late Triassic bivalves from Sonora, Mexico, figures 5.6-5.9.
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«Affinities: The most similar species is W. bodenbenderi (BEHR.), which is also described here and which has a deeper byssal sinus and su bequal auricles. The apical angle is larger in W. bodenbenderi which also has fewer and different auricular crura, not parallel to the dorsal margin but to the inner margin of the auricles. The ornamentation is also quite different; on the right valve the ribs are less prominent, wider and they are frequently split ot have minor intercalated ribs, especially on the antetior and posterior portions of the shell. On the left valve the difference in ornamentation is even greater; in W. bodenbenderi the radial ribs are flat-topped, convex in transverse section, and at least as wide as the interval between them. They can also be subdivided or have minor intercalated ribs, especially on the flanks (see BEHRENDSEN 1891, Taf. 22, Fig. 3; 1922, lám. 1, fig. 10; BURCKHARDT 1902, pI. 4, fig. 1-2; JAWORSKI 1914a, fig. 9-10; 1925b, lám. 1, fig. 1; 1926, text-fig. 2 as Vola simplex; WEAVER 1931, pI. 33, fig. 184; LEANZA 1942b, lám. 7, fig. 6; lám. 8, fig. 1-2; and here on pI. 10, 11 and 12).
Weyla titan (MÖRICKE), which may be considered as a subspecies of W. alata, is known from some Chilean localities (MÖRICKE 1894: 41; R. PHILIPPI 1899: 29, lám. 15) and it was also mentioned from Peru (BRAVO 1906: 106-108) and Argentina (JAWORSKl 1914a: 283-284; 1915: 439; 1925b: 53). This shell is much larger, posteriorly elongated and the ribs become flat and wider than the interspaces near the ventral margin. Weyla alata angustecostata (PHILIPPI) has narrower ribs which have a completely flat top, the intervals between the ribs are wider and the inner surface of the shell is smooth except near the ventral margin (text-fig. 23). The species W. rollieri (COSSMANN), from the early Pliensbachian of France (COSSMANN 1916, pI. 5, fig. 13-15; DUBAR 1925, fig. 41-42, pI. 5, fig. 10-12) differs in having fewer radial ribs (8 to 10), a different cross section of the right valve ribs and larger and more pointed auricles.
W. alata shares with W. mexicana (JAWORSKI), from the early Jurassic of Mexico and Argentina (JAWORSKI 1929: 2-5, Taf. 1, Fig. 1a-c, 2, 3; BURCKHARDT 1930: 23; KING 1939: 1654) the form and relative convexity of both valves. It has the same number of radial ribs and a pair of radial ribs on the left valve auricles, but W. mexicana has subequal ribs and intervals that are triangular in transverse section on both valves. This species also has a peculiar concentric ornamentation of strong V-shaped growth lamellae, particularly developed on the interspaces. W. ambongoensis (THEVENIN), known from the Toarcian of Malagasy (THEVENIN 1908: 24, pI. 4, fig. 2-3; DUBAR 1948: 220; ARKELL 1956: 342; BESAIRE & COLLIGNON 1956: 58), Morocco (DUBAR 1948: 220, pI. 29, fig. 7-9), Pakistan (HOLLAND 1909: 27-28; ARKELL 1956: 398), Kenya (ARKELL 1956: 318; COX 1965: 59, pI. 7, fig. 9; see also text-fig. 20 here), Saudi Arabia (ARKELL et a1. 1952; ARKELL 1956: 300) and China (WEN et a1. 1976: 101, pI. 24, fig. 6-9) also has subtriangular ribs, but differs from all other Weyla species because its shell, originally concave-convex, becomes biconvex in adults and never exceeds 5 em in length. The radial ribs of W. ambongoensis, up to 12 in number, may bear tubercles or spine bases on both valves (see text-fig. 20). Other species that also have V-shaped ribs on both valves are the European W. pradoana (DE VERNEUIL & COLLOMB 1853) and W. lacazei (HAIME 1855, see also CALZADA 1983, fig. 3: 1a-c and MOUTERDE et al. 1983, pI. 13, fig. 6). A pair of very closely related (or even possibly synonymous) species were described from Morocco and Spain: W. ayarti (DUBAR 1948: 217, pI. 28, fig. 26; ARSICAULT 1965: 41) and W. almelae QUINTERO & DE LA REVILLA (1966: 41, lam. 5, fig. la-b), but these shells are smaller than W. alata and only have up to six radial ribs on each valve. The left valve of these species is unknown.
Other species, here grouped in the new subgenus Lywea, can be easily distinguished from W. alata by having both valves convex and V-shaped ribs and interspaces. "Parapecten" ntlakapamuxanus CRICKMAY (1930, pI. 5, fig. f-g), from the Early Jurassic of British Columbia, Canada, and "Pecten" argentarius GABB (1877: 293-294, pI. 41, fig. 12), from the Pliensbachian (?) of Perú, are both imperfectly known species, with which it is difficult to establish comparisons. Remarks: W. alata alata was described by VON BUCH (1838: 55-56) and first figured one year later by the same author (1839: 3-4, 20, pI. 1, fig. 1-4) on the basis of material collected by HUMBOLDT in Perú, originally referred to the early Cretaceous. VON BUCH'S figure represents a somewhat inequilateral specimen, a common feature in distorted Weyla specimens. Nevertheless this author assigned special importance to the fact that the antetior portion of the shell extends in a wing-like fashion (therefore the name "alata"). This character was widely discussed later and while some authors (such as BAYLE & COQUAND 1851; JAWORSKI 1914a) considered that the figured specimen was simply an individual variant within a species with equilateral shells, others (such as GIEBEL in BURMEISTER & GIEBEL 1861; R. PHILIPPI 1899) maintained that it is valid as a character and typical of the species. VON BUCH'S figure was reproduced by R. PHILIPPI (1899) and incorrectly interpreted by JAWORSKI (1914a) as a variety of W. alata, for which he proposed the name Vola alata var. aspera.
Only few years after VON BUCH'S description of W. alata, D'ORBIGNY (1843: 106, pI. 22, fig. 5-9), on the basis of material collected by DOMEYKO in Chile, proposed the new name Pecten dufrenoyi, also referring it to the early Cretaceous. This name is here regarded as a junior synonym of W. alata (V. BUCH). This same opinion was given by BAYLE & COQUAND (1851), who had examined D'ORBIGNY'S specimens and is here additionally supported by the personal observation of specimens collected and mentioned by DARWIN (1846), determined as Pecten Dufrenoyi by D'ORBIGNY himself. These specimens are housed at the BMNH and show all the features of the species here described (see text-fig. 18, 1-2). The unusual ribbing of the right valve that appears in D'ORBIGNY'S figure could not be seen on any of the numerous specimens of Weyla from Chile examined in different museum collections. As BURMEISTER & GIEBEL (1861) already noticed, it is probable that the drawing was a reconstruction from incomplete specimens based on the assumption that the ribbing was strictly symmetrical on both valves. This synonymy was accepted by most authors, except HUPÉ (1854) and BEHRENDSEN (1891). The possibility that D'ORBlGNY'S specimen belongs to W. alata angustecostata (PHILIPPI) instead, cannot be completely dismissed and in that circumstance, D'ORBIGNY'S name should be used for this subspecies of W. alata. BAYLE & COQUAND (1851) excelently illustrated W. alata and settled the age problem, dating it as early Jurassic. All other authors followed their interpretation of the taxon. R. PHILIPPI (1899) reproduced their illustration and proposed for it the new name Pecten commutatus, which is not only unnecessary but also a junior homonym of P. commutatus MONTEROSATO 1875, and thus invalid. The first earlyJurassic fossil mentioned for Argentina was also referred to Pecten alatus, mentioned by STROBEL (1870, 1875) from the Portezuelo Ancho region. Unfortunately there are not enough data to establish if this record corresponds to W. alata or to the species from the same place later described by BEHRENDSEN (1891) as W. bodenbenderi, since both species occur near that locality. W. alata was very soon regarded as a characteristic Andean early Jurassic species and as early as 1878 GOTTSCHE provided a summary of its geographical distribution. Some of the numerous species described by R. PHILIPPI (1899) are here tentatively included as synonyms of W. alata, pending a revision of the Chilean material. This species was frequently mentioned for several South American localities and was figured by JAWORSKI (1914a, figures reproduced in HOLDER 1964 and in COX et al. 1969), STEINMANN (1929, figures reproduced in COX et al. 1969), WEAVER (1931), GEYER (1973), RANGEL (1978), HILLEBRANDT (1980) and PÉREZ (1982). SCHLAGINTWEIT (1912) doubted of the early Jurassic age of at least part of VON BUCH'S material, especially that collected between Guambos and Montan, where according to SCRLAGINTWEIT there are no outcrops of early Jurassic age. On the other hand early Cretaceous beds do crop out there and could bear bivalves referable to Neithea. This raises the question of the real identity of the species, because it is probable that the rest of the specimens that VON BUCH examined (i. e. from Huancavélica) are in fact of early Jurassic age. The designation of a lectotype to stabilize the concept of Pecten alatus VON BUCH (and consequently the genus that it typifies) is thus of fundamental importance, but cannot he done without the observation of VON BUCH'S specimens, if they are still available.» DAMBORENEA, S. E. 1987. Early Jurassic Bivalvia of Argentina; Part 2, Superfamilies Pteriacea, Buchiacea and part of Pectinacea. Palaeontographica, Abteilung, A, Palaeozoologie-Stratigraphie, 199 (4-6): 113-216, pls. 1-14. [p. 174-177]
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Weyla (Weyla) alata alata (von Buch, 1838); S. E. Damborenea, 1987, Early Jurassic Bivalvia of Argentina, Part 2, Superfamilies Pteriacea, Buchiacea and part of Pectinacea, plate 9, figure 1; plate 10, figures 1-5.
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«Remarks. This species is abundant in the Lias of Neuquen and Mendoza, but usually the specimens are greatly weathered, causing apparent variations in the character of the ornamentation. Seldom are specimens obtained in as good a state of preservation as those in the drawings of Bayle et Coquand (14), pl. 5, figs. 1 and 2. The immature specimens have been identified as Pecten pradoanus Vern et ColI (18) Behrendsen, p. 391, pl. 22, figs. 1a-d, but exhibit the characteristics of the umbonal portions of the larger and more mature individuals. This condition was earlier recognized by Moericke (262), p. 39, who placed Behrendsen's form in synonymy with V. alata. Jaworski in his work (190) on the anatomy of the hard parts of the Liassic Volas of South America has aided greatly in clearing up the distinctions between the different species which have been created from time to time to accommodate the differences in shell outline and sculpture so common among the South American specimens. Ten distinct species have been described by Philippi (300) in his work on the Mesozoic fossils of Chile, but many of the illustrations are from specimens in a poor state of preservation, and probably when a revised study of his material is made some will be found to be synonymous with V. alata or V. bodenbenderi as has already been suggested by Jaworski (190), p. 276, in the case of V. curvicosta.
V. alata (Von Buch) Bayle et Coquand differs from V. bodenbenderi in having the right valve more convex, the radiating ribs broader and higher, the interspaces narrower, and with the presence of a well developed minor riblet traversing the center of each interspace, and usually two auxiliary smaller riblets on either side where the major ribs and interspaces join. Jaworski (196), p. 163, pl. 1, fig. 1, has recently described Vola simplex from the lower Lias of Portezuelo Ancho in Southern Mendoza, which is characterized by having 12 prominent, rounded, radiating ribs in the left valve, with broad interspaces, each of which contains a single, fairly well developed, minor, radiating rib. These ribs gradually disappear toward the umbonal region. In addition there are occasionally fine radiating striae in contrast to V. alata, where there exist broad interspaces of the left valve without the single median minor rib. Moericke has grouped those very large types of V. alata (262), p. 39, in which the ribs widen and flatten as they approach the basal margin so as to become 3 times as wide as the interspaces, into the variety V. alata var. titan.» WEAVER, C. E. 1931. Paleontology of the Jurassic and Cretaceous of West Central Argentina. Memoirs of the University of Washington, 1: 1-469, pls. 1-62. [p. 282]
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Vola alata (Von Buch) Bayle et Coquand; C. E. Weaver, 1931, Paleontology of the Jurassic and Cretaceous of West Central Argentina, plate 31, figure 179; plate 32, figure 181.
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«Décrite par M. de Buch sous le nom de Pecten alatus, cette espèce a reçu de M. Alc. d'Orbigny celui de Pecten Dufrenoyi. Nous avons pu nous assurer, en comparant entre eux un grand nombre d'individus et ceux même qui ont servi aux descriptions de M. d'Orbigny, que ces deux espèces doivent être réunies, la figure donnée par M. de Buch ne constituant qu'une simple variété.
Le faciès crétacé de cette espèce a fait penser à M. d'Orbigny qu'elle caractérisait dans le Chili l'étage de la craie chloritée; ce qui démontre le danger de tirer des conclusions àpriori d'après telle ou telle forme de genres, pour les appliquer à la classification des terrains. Le Pecten alatus a été rapporté par M. de Humboldt des environs de Lima; il a été retrouvé à Guanca-Velica, à Copiapo, sous le 26° degré de latitude S., et par M. Domeyko, à Mandas, à Tres-Cruces et à Jorquera. Il paraît être fort abondant. La collection de l'École des mines en possède une très belle suite de tous les âges.» BAYLE, E. & H. COQUAND. 1851. Mémoire sur les fossiles recueillis dans le Chili par M. Ignace Domeyko et sur les terrains auxquels ils appartiennent. Mémoires de la Société Géologique de France [2me Série], 4: 1-47, pls. 1-8. [p. 15]
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Pecten alatus, De Buch; E. Bayle & H. Coquand, 1851, Mémoire sur les fossiles recueillis dans le Chili par M. Ignace Domeyko, plate 5, figures 1, 2.
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